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subgenera

Entoloma Nolanea Leptonia Pouzarella Clitopiloides Allocybe Alboleptonia Omphaliopsis ParaleptoniaClaudopus


E. prunuloides E. bloxamii

E. sinuatum

E. rubellummoserianum
E. moserianumclypeatum
E. clypeatumniphoides
E. niphoidessaundersii
E. saundersiisepium
E. sepiumrhodopolium
E. rhodopoliumnoordeloosii
E. noordeloosiimyrmecophilum
E. myrmecophilumgriseoluridum
E. griseoluridumgriseopruinatum
E. griseopruinatumlividoalbum
E. lividoalbumpolitum
E. politumcaccabusE. caccabussphagneti
E. sphagnetianthracinellum
E. anthracinellumbipelleE. bipellepallideradicatum
E. pallideradicatumturbidumE. turbidum

Classification: Subgenus Entoloma

Nolanidea / Rhodopolia/ Polita/ Turfosa/ Sphagnophila/ Pseudonolanea/Pallideradicata

Fruitbodies tricholomatoid and fleshy (sections Entoloma, Nolanidea, Rhodopolia and Sphagnophila, Pallideradicata , less often collybioid or omphalioid (sections Polita and Turfosa). Pileus smooth, hygrophanous or not. Pileipellis a cutis or ixocutis of narrow, 2-10 µm wide hyphae with intracellular or incrusted pigment. Cystidia exceptional, and if cheilocystidia present, they usually are of a simple basidioliform type. Hyphae of trama of short elements (e.g. 40-100 x 5-15 µm). Clamp-connections usually abundant in all tissues.

In Europe this subgenus is represented with seven sections:

Section Entoloma, subsect. Entoloma
Habit tricholomatoid, fleshy. Pileus not hygrophanous, not or indistinctly translucently striate; occurring in autumn. Spores small, isodiametrical, often weakly and many-angled.
Type species: Entoloma prunuloides (Fr.: Fr.) Quél.


1.   Entoloma prunuloides (Fr.: Fr.) Quél.
1a. Entoloma prunuloides var. prunuloides
1b. Entoloma prunuloides var. obscurum Arnolds & Noordel.
2.   Entoloma myochroum Noordel. & E. Ludw.
3.   Entoloma bloxamii (Berk. & Broome) Sacc.
3a. Entoloma bloxamii f. bloxamii
3b. Entoloma bloxamii f. caesiolamellatum Noordel. & Wölfel
4.   Entoloma nitidum Quél.
5.   Entoloma rubellum (Scop.) Gillet
6.   Entoloma viridans (Fr.) P. Karst.
7.   Entoloma alcedicolor Arnolds & Noordel.

Section Entoloma, subsect. Sinuata
Spores iso- to heterodiametrical, 5-7 (-8) angled in side-view.
Type species: Entoloma sinuatum (Bull. ex Pers.) P. Kumm.


8.   Entoloma sinuatum (Bull. ex Pers.) P. Kumm.
9.   Entoloma moserianum Noordel.
10. Entoloma luteobasis Ebert & E. Ludw.
11. Entoloma olidum Noordel.

 

Section Nolanidea
This section essentially is created for a group of similar species sharing a particular ecology. They grow all together with Rosaceae or Ulmaceae, forming a special type of ectomycorrhiza with members of these families. In Europe, they all appear from late winter to spring. Their morphology is more or less intermediate between sect. Entoloma and the rhodopolium-group. Some taxa have a non-hygrophanous pileus with aeriferous-fibrillose patches (E. saundersii, to some extent also E. sepium), others have a glabrous, hygrophanous pileus reminiscent E. rhodopolium and relatives.
Type-species: Entoloma clypeatum (L) P. Kumm.


12.   Entoloma clypeatum (L.) P. Kumm.
12a. Entoloma clypeatum var. clypeatum
12b. Entoloma clypeatum f. pallidogriseum Noordel.
12c. Entoloma clypeatum f. xanthophyllum Noordel.
12d. Entoloma clypeatum var. defibulatum Noordel.
12e. Entoloma clypeatum var. hybridum Romagn.
13.   Entoloma aprile (Britzelm.) Sacc.
14.   Entoloma niphoides Romagn. ex Noordel.
15.   Entoloma sepium (Noul. & Dass.) Richon & Roze
16.   Entoloma saundersii (Fr.) Sacc

Section Rhodopolia. Pileus distinctly hygrophanous, in thin-fleshed species usually also distinctly translucently striate. Pigment intracellular, incrusting or a combination of both types
Type-species: Entoloma rhodopolium (Fr.: Fr.) P. Kumm.


17.   Entoloma rhodopolium (Fr.: Fr.) P. Kumm.
17a. Entoloma rhodopolium var. rhodopolium
17b. Entoloma rhodopolium var. nidorosum (Fr.) Noordel.
17c. Entoloma rhodopolium var. pseudopolitum Noordel.
19.   Entoloma leucocarpum Noordel.
20.   Entoloma roseoalbum Arnolds & Noordel.
21.   Entoloma pseudoexcentricum (Romagn.) Kreisel
21a. Entoloma pseudoexcentricum var. pseudoexcentricum
21b. Entoloma pseudoexcentricum var. microsporum Noordel. & E. Ludw.
22.   Entoloma noordeloosi Hauskn.
23.   Entoloma rubrobasis Noordel.
24.   Entoloma difforme Naveau.25. Entoloma sericatum (Britzelm.) Sacc.
25a. Entoloma sericatum var. sericatum
25b. Entoloma sericatum var. saliciphilum (Noordel.) Noordel.
26.   Entoloma majaloides P.D. Orton
27.   Entoloma sordidulum (Kühner & Romagn.) P.D. Orton
28.   Entoloma subradiatum (Kühner & Romagn.) Mos.
29.   Entoloma inusitatum Noordel., Enderle & H. Lammers 30.   Entoloma myrmecophilum (Romagn.) Mos.
30a. Entoloma myrmecophilum var. myrmecophilum
30b. Entoloma myrmecophilum var. coalescens Noordel. & Luhmann
31.   Entoloma venosum Gillet
32.   Entoloma gerriae Noordel.
33.   Entoloma griseorugulosum Noordel. & Fern. Sas.
34.   Entoloma lividoalbum (Kühner & Romagn.) Kubicka
35.   Entoloma griseoluridum (Kühner) Mos.
36.   Entoloma griseopruinatum Noordel. & Cheype
37.   Entoloma singulare (Romagn.) Bon & Courtec.
38.   Entoloma brassicolens (D.A. Reid) Noordel.
39.   Entoloma transvenosum Noordel.
39a, Entoloma transvenosum var. transvenosum
39b. Entoloma transvenosum var. microsporum Noordel. & Hauskn.
40.   Entoloma kallioi Noordel.

Section Polita. Habit collybioid, omphalioid, rarrely tricholomatoid; pigment intracellular
Type-species: Entoloma politum (Pers.: Fr.) Donk

41.  Entoloma politum (Pers.: Fr.) Donk
42.  Entoloma caccabus (Kühner) Noordel.
43.   Entoloma bisporigerum (P.D. Orton) Noordel.
44.   Entoloma engadium (Horak) Noordel.
45.   Entoloma anthracinum (Favre) Noordel.
46.   Entoloma melenosmum Noordel.
47.   Entoloma cistophilum Trimbach
48.   Entoloma caeruleopolitum Noordel. & Brandt?Petersen
49.   Entoloma reaae (Maire) Noordel.
50.   Entoloma nausiosme Noordel.
51.   Entoloma fridolfingense Noordel. & Lohmeyer

Section Sphagnophila. Habit tricholomatoid; spores many-angled, almost nodulose, pileus covered when young with fibrillose patches of what may be originating from universal veil; pigment abundant, brown, intracellular; clamp-connections present. In peat bogs among dead or living Sphagnum.
Type species: Entoloma sphagneti Naveau.

52. Entoloma sphagneti Naveau

Section Pseudonolanea. Basidiocarps mycenoid or collybioid, rarely tricholomatoid; often resembling a Nolanea species like E. sericeum, but trama typically entolomatoid with short, sausage-shaped elements. Clamp-connections usually present in all tissues.
Type-species: Entoloma alpicola (Favre) Noordeloos)

53a. Entoloma atrosericeum var. atrosericeum
53b. Entoloma atrosericeum var. fusipes Noordel.
54.   Entoloma bipelle Noordel. & Borgen
55.   Entoloma borgenii Noordel.
56.   Entoloma svalbardense
57.   Entoloma subarcticum Noordel.
58.   Entoloma alnobetulae (Kühner) Noordel.
59.   Entoloma alpicola (Favre) Noordel.
60.   Entoloma subsepiaceum (Kühner) Noordel.
61.   Entoloma rimulosum Noordel.
62.   Entoloma bicorne Noordel.
63.   Entoloma anthracinellum (M. Lange) Noordel.
64.   Entoloma philocistis Hauskn. & Noordel.
65.   Entoloma atroenigmaticum Noordel. & Hauskn.
66.   Entoloma vindobonense Noordel. & Hauskn.
67.   Entoloma valdeumbonatum Noordel. & Meusers

Section Pallideradicata. Habit (slender) tricholomatoid with conical to convex, umbonate pileus, adnate to almost free lamellae, and relatively slender stipe. Pileus not or slightly hygrophanous, not translucently striate, glabrous to lustrous-fibrillose or very minutely radially veined, white or very pale coloured. Spores heterodiametrical. Lamella edge fertile or heterogeneous. Cheilocystidia present or absent. Pleurocystidia absent. Pileipellis a well differentiated (ixo-)cutis of narrow cylindrical hyphae with intracellular and/or parietal pigment; trama composed of relatively long hyphae. Clamp-connections absent.
Type species: Entoloma pallideradicatum Noordel. & Hauskn.

68. Entoloma pallideradicatum Noordel. & Hauskn.
69.   Entoloma eximium (Romagn.) Noordel.


Section Turfosa The species of this section have in common a very peculiar spore type, with many angles and very thin walls compared with other Entoloma species. The spores stain well in cotton-blue (cyanophilous).
Type species: Entoloma turbidum (Fr.: Fr.) Quél.


70.   Entoloma turbidum (Fr.: Fr.) Quél.
71.   Entoloma vinaceum (Scop.) Arnolds & Noordel.
71a. Entoloma vinaceum var. vinaceum
71b. Entoloma vinaceum var. fumosipes Arnolds & Noordel.
71c. Entoloma vinaceum var. violeipes Arnolds & Noordel.
72.   Entoloma pseudoturbidum (Romagn.) Mos.
73.   Entoloma pluteisimilis Noordel. & C.E. Hermos.
74.   Entoloma cancrinellum (M. Lange) Noordel.

Species of uncertain affinity.

75.   Entoloma lactarioides Noordel. & Liiv.